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Vision is the perception of objects based  on the light that they emit or reflect.   00:03
Human eyes can only detect visible light -  a narrow range of electromagnetic radiation,   00:08
roughly from 400 to 750 nm in wavelengths.  00:14
The eye consists of optical components, which  work like a camera, capturing and focusing images;   00:19
and neural components that convert these images  into nerve impulses and send them to the brain.  00:26
The main optical components are the cornea,  the lens, and the iris. The cornea and the   00:32
lens refract light and focus the image on  the retina. The iris acts as an aperture,   00:38
it controls the amount of light that enters  the eye by adjusting the size of the pupil.  00:45
The neural components are the retina - a  light-sensitive tissue lining the inner   00:50
surface of the eye, and the optic nerve. Light  is absorbed by photoreceptor cells in the retina.   00:54
The optical information is then passed through  several cell layers, where it is converted into   01:01
action potentials and sent, via the optic  nerve, to the visual cortex of the brain.  01:07
The fovea is the central part of the retina  where the sharpest central vision is achievable.  01:13
The optic disk, where the optic nerve  leaves the eye, has no photoreceptor cells.   01:19
It corresponds to the blind spot in the  visual field. If an object falls on that spot,   01:25
it would generate no visual information. However,  instead of leaving a black hole in the vision,   01:31
the brain fills it in with visual  information from around the object.  01:37
The major photoreceptor cells of the retina  are rods and cones. Rod cells are responsible   01:41
for night vision. They can detect dim light,  but provide low-resolution images and cannot   01:48
differentiate colors. Cones function in bright  day light. They detect colors, and provide high   01:53
resolution details. There are 3 kinds of cones  named after the color that they absorb best:   02:01
red, green and blue. A color is perceived based  on proportions of signals coming from these cones.   02:07
Color blindness occurs when a person  lacks a certain kind of cones.  02:15
The ability of photoreceptor cells to detect light   02:20
is due to their light-receptor  molecules, called visual pigments.   02:24
It’s rhodopsin in rods, and iodopsins in  cones. These molecules consist of 2 components:   02:28
a protein called opsin, and a vitamin A-derivative  called retinal. The retinal component is   02:36
identical for all visual pigments, but the opsin  is different for rods and each type of cones.   02:43
Different opsins absorb different wavelengths,  allowing detection of different colors.  02:50
In the dark, there is a so-called  dark current in photoreceptor cells.   02:56
This is due to the presence of cGMP, which  permits a constant influx of sodium. The cells are   03:01
depolarized, they release the neurotransmitter  glutamate at the synapse with bipolar cells.  03:08
The retinal exists in 2 conformations:  cis and trans. In the dark,   03:15
the cis-form is bound to opsin, keeping  it inactive. As the retinal absorbs light,   03:21
it changes to trans-form and dissociates from the  opsin, which now becomes an active enzyme. The   03:28
enzyme degrades cGMP, sodium channel closes, dark  current stops and so does glutamate secretion.   03:35
The drop in glutamate tells the bipolar  cells that light has been absorbed.   03:44
The information is then transmitted  to ganglion cells - the only cells   03:49
within the retina that generate action  potentials and send them to the brain.  03:53
On average, each ganglion cell receives  signals from over a hundred of rods. This   03:59
degree of convergence is at the basis of the high  sensitivity of rod cells. A dim light produces   04:04
only a weak signal in a rod, but together,  hundreds of these signals converge and become   04:11
one strong signal acting on a single ganglion  cell. However, as the signal comes from a large   04:17
area of the retina, the image resolution is poor. The cones have a much lower degree of convergence.   04:23
The fovea in particular has only cones and  no rods, and each cone conveys signal to one   04:31
ganglion cell. Because one ganglion cell receives  input from a very small area of the retina, this   04:38
setup produces high resolution images. But high  resolution comes with low sensitivity, because   04:44
each cone must be stimulated with a signal  strong enough to generate action potentials   04:50
in the ganglion cell. This also explains  why there is no color vision in dim light.  04:56
There exist other cell types that form  connections between photoreceptor cells,   05:03
or bipolar cells. They detect changes in light  intensity within an image and provide additional   05:08
information about contrast and edges of objects. Some of the ganglion cells also absorb light   05:14
directly, but not for the purpose of forming  images. They transmit information about light   05:22
intensity to the brainstem, as part of the reflex  that controls pupil size; and to the hypothalamus,   05:28
as input for the sleep-wake cycle. The bipolar cells are first-order neurons,   05:35
and ganglion cells are second-order neurons. The  axons of ganglion cells form the optic nerve.   05:41
The 2 optic nerves from the 2  eyes converge at the optic chiasm.   05:48
Here, the medial half of nerve fibers from  each eye cross to the other side of the brain.   05:53
Most of the fibers then continue to the thalamus  and synapse with third-order neurons, whose axons   05:59
project to the primary visual cortex. Some fibers  take a different route: they terminate in the   06:05
midbrain and are responsible for pupillary light  reflex and accommodation reflex, among others.  06:12
Note that objects in the left visual field  are perceived by the right side of the brain,   06:19
which also controls motor responses of the  body’s left side – the same side as the objects. 06:24

– English Lyrics

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Lyrics & Translation

[English]
Vision is the perception of objects based  on the light that they emit or reflect.  
Human eyes can only detect visible light -  a narrow range of electromagnetic radiation,  
roughly from 400 to 750 nm in wavelengths. 
The eye consists of optical components, which  work like a camera, capturing and focusing images;  
and neural components that convert these images  into nerve impulses and send them to the brain. 
The main optical components are the cornea,  the lens, and the iris. The cornea and the  
lens refract light and focus the image on  the retina. The iris acts as an aperture,  
it controls the amount of light that enters  the eye by adjusting the size of the pupil. 
The neural components are the retina - a  light-sensitive tissue lining the inner  
surface of the eye, and the optic nerve. Light  is absorbed by photoreceptor cells in the retina.  
The optical information is then passed through  several cell layers, where it is converted into  
action potentials and sent, via the optic  nerve, to the visual cortex of the brain. 
The fovea is the central part of the retina  where the sharpest central vision is achievable. 
The optic disk, where the optic nerve  leaves the eye, has no photoreceptor cells.  
It corresponds to the blind spot in the  visual field. If an object falls on that spot,  
it would generate no visual information. However,  instead of leaving a black hole in the vision,  
the brain fills it in with visual  information from around the object. 
The major photoreceptor cells of the retina  are rods and cones. Rod cells are responsible  
for night vision. They can detect dim light,  but provide low-resolution images and cannot  
differentiate colors. Cones function in bright  day light. They detect colors, and provide high  
resolution details. There are 3 kinds of cones  named after the color that they absorb best:  
red, green and blue. A color is perceived based  on proportions of signals coming from these cones.  
Color blindness occurs when a person  lacks a certain kind of cones. 
The ability of photoreceptor cells to detect light  
is due to their light-receptor  molecules, called visual pigments.  
It’s rhodopsin in rods, and iodopsins in  cones. These molecules consist of 2 components:  
a protein called opsin, and a vitamin A-derivative  called retinal. The retinal component is  
identical for all visual pigments, but the opsin  is different for rods and each type of cones.  
Different opsins absorb different wavelengths,  allowing detection of different colors. 
In the dark, there is a so-called  dark current in photoreceptor cells.  
This is due to the presence of cGMP, which  permits a constant influx of sodium. The cells are  
depolarized, they release the neurotransmitter  glutamate at the synapse with bipolar cells. 
The retinal exists in 2 conformations:  cis and trans. In the dark,  
the cis-form is bound to opsin, keeping  it inactive. As the retinal absorbs light,  
it changes to trans-form and dissociates from the  opsin, which now becomes an active enzyme. The  
enzyme degrades cGMP, sodium channel closes, dark  current stops and so does glutamate secretion.  
The drop in glutamate tells the bipolar  cells that light has been absorbed.  
The information is then transmitted  to ganglion cells - the only cells  
within the retina that generate action  potentials and send them to the brain. 
On average, each ganglion cell receives  signals from over a hundred of rods. This  
degree of convergence is at the basis of the high  sensitivity of rod cells. A dim light produces  
only a weak signal in a rod, but together,  hundreds of these signals converge and become  
one strong signal acting on a single ganglion  cell. However, as the signal comes from a large  
area of the retina, the image resolution is poor. The cones have a much lower degree of convergence.  
The fovea in particular has only cones and  no rods, and each cone conveys signal to one  
ganglion cell. Because one ganglion cell receives  input from a very small area of the retina, this  
setup produces high resolution images. But high  resolution comes with low sensitivity, because  
each cone must be stimulated with a signal  strong enough to generate action potentials  
in the ganglion cell. This also explains  why there is no color vision in dim light. 
There exist other cell types that form  connections between photoreceptor cells,  
or bipolar cells. They detect changes in light  intensity within an image and provide additional  
information about contrast and edges of objects. Some of the ganglion cells also absorb light  
directly, but not for the purpose of forming  images. They transmit information about light  
intensity to the brainstem, as part of the reflex  that controls pupil size; and to the hypothalamus,  
as input for the sleep-wake cycle. The bipolar cells are first-order neurons,  
and ganglion cells are second-order neurons. The  axons of ganglion cells form the optic nerve.  
The 2 optic nerves from the 2  eyes converge at the optic chiasm.  
Here, the medial half of nerve fibers from  each eye cross to the other side of the brain.  
Most of the fibers then continue to the thalamus  and synapse with third-order neurons, whose axons  
project to the primary visual cortex. Some fibers  take a different route: they terminate in the  
midbrain and are responsible for pupillary light  reflex and accommodation reflex, among others. 
Note that objects in the left visual field  are perceived by the right side of the brain,  
which also controls motor responses of the  body’s left side – the same side as the objects.

Key Vocabulary

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Vocabulary Meanings

vision

/ˈvɪʒən/

B1
  • noun
  • - the ability to see; sight

light

/laɪt/

A1
  • noun
  • - the natural agent that stimulates sight and makes things visible
  • noun
  • - low-powered
  • verb
  • - to illuminate

eye

/aɪ/

A1
  • noun
  • - the organ of sight

detect

/dɪˈtɛkt/

B1
  • verb
  • - to discover or identify the presence or existence of

visible

/ˈvɪzəbl̩/

B1
  • adjective
  • - able to be seen

radiation

/ˌreɪ.diˈeɪ.ʃən/

B2
  • noun
  • - energy emitted in the form of waves or particles

wavelength

/ˈweɪv.lɛŋkθ/

C1
  • noun
  • - the distance between successive crests of a wave

cornea

/ˈkɔːr.ni.ə/

C2
  • noun
  • - the transparent layer forming the front of the eye

lens

/lɛnz/

B2
  • noun
  • - a transparent optical component that focuses light

iris

/ˈaɪrɪs/

C1
  • noun
  • - part of the eye controlling pupil size
  • noun
  • - a flower

pupil

/ˈpjuːpl/

B1
  • noun
  • - the opening in the eye allowing light entry
  • noun
  • - a student

retina

/ˈrɛtənə/

C2
  • noun
  • - the inner layer of the eye containing photoreceptor cells

optic

/ˈɒptɪk/

C1
  • adjective
  • - relating to sight or the eye

nerve

/nɜːrv/

B1
  • noun
  • - a bundle of fibers that transmits impulses

cell

/sɛl/

A2
  • noun
  • - the smallest structural and functional unit of an organism

rod

/rɒd/

C1
  • noun
  • - a type of photoreceptor cell for low-light vision

cone

/koʊn/

C1
  • noun
  • - a type of photoreceptor cell for color vision

brain

/breɪn/

A2
  • noun
  • - the organ of the central nervous system

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